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Proof: This is immediately established by noting that, when p1 and p2 are constant, the expected lifespan of A is 1/p2 and the expected number of offspring is simply the number of offspring expected during the expected lifespan, i.e., p1/p2. In more detail, the probability of A surviving for exactly T time-steps is p(T)= (1 - p2)T-1 ·p2, and the expected number of ''offspring" during that interval is C0110-04.gif. Thus, the expected number of "offspring" during A's lifespan is
C0110-01.gif
But C0110-05.gif converges to (1/p2)2 (as may be easily verified by taking the derivative of both sides of the identity (1/1 - x) = 1 + x + x2 + . . . ). Therefore
C0110-02.gif
For plans in C0110-06.gif the interpretation of this lemma is direct: The probability of Ah being selected to produce an offspring A' during time t is C0110-08.gif where C0110-07.gif, while the probability of Ah being deleted at the end of that time-step is 1/M. Hence, if C0110-13.gifchanges negligibly over Ah's lifespan, the expected number of offspring is
C0110-03.gif
C0110-14.gif can be looked upon as a "normalized" payoff, the "usefulness" of Ahbeing measured relative to the average performance of the other members in the population. With this arrangement the expected number of offspring of Ah is greater than 1 just in case Ah's performance is above average. Since C0110-09.gif is not stationary for plans C0110-10.gif, the probability p1 does not in fact remain constant (though, over the expected lifespan of a structure, it will not often change greatly). If C0110-09.gif increases (as it will generally with a good plan), then Ah will receive fewer offspring than predicted by the calculation of p1 at the time Ah originated. That is, the performance of Ah looks less promising relative to the current average, so trials of Ah are curtailed. If C0110-09.gif decreases, the opposite effect occurs. Still, the expected number of offspring varies in direct relation to Ah's relative performance, so that plans in C0110-06.gif satisfy the (informal) characterization of reproductive plans.
A slight change in the form of the algorithms in C0110-06.gif yields a class of algorithms Rd wherein a time-step is a "generation" during which each individual C0110-15.gifis replaced, deterministically instead of as an expectation, by C0110-12.gif offspring. Thus, for Rd, C0110-11.gif consists of the set of all offspring of the individuals in C0177-03.gif. (To keep the population level at M individuals a special kind of rounding

 
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